Russula Pers.
Russula Pers. is a genus of high species diversity with a comprehensive wide distribution from frigid to tropical forests (Kundsen and Borgen 1982; Singer 1986; Buyck 1989; Buyck et al. 1996; Miller et al. 2012). Russula is evidenced from ITS, nLSU and rpb2 to be a monophyletic genus (Buyck et al. 2008, 2010), but it contains stipitate epigeous, hypogeous, and pleurotoidformed fruiting bodies (Buyck and Hoyak 1999; Miller et al. 2001; Larsson and Larsson 2003; Lebel and Tonkin 2007). Nine subgenera have been introduced in Russula based on morphological characteristics, such as taste of fruiting bodies, colour of spore print, shape of pileipellis hyphal extremities, existence of lamellulae, dermatocystidia and primordial hyphae (Romagnesi 1967, 1985, 1987), and phylogenetic data is needed in classification (Eberhardt 2002; Li and Wen 2009; Li 2014; Li et al. 2015a, b). A total of 22 Russula taxa have been described from China and the adjacent Himalayan Mountain in recent years (Das et al. 2005, 2006a, b, 2010, 2013, 2014; Wang et al. 2009b; Li et al. 2011, 2012, 2013a, b, 2015a, b, d). Two taxa are newly described from Tibet Plateau based on morphological characters and phylogenetic analyses. The phylogenetic tree for Russula is presented in Fig. 1.

Fig. 1 Phylogram generated from maximum parsimony (PAUP* v.4.01) analysis based on ITS sequence data of Russula. Sequences used in this study have been sampled from previous studies to represent the major phylogenetic clades of Russula (Eberhardt 2002; Miller and Buyck 2002). Maximum parsimony bootstrap support values above 50 % and Bayesian posterior probabilities greater than 0.9 are indicated above or below the nodes (BS/PP), new taxa are shown in blue. Holotype are shown in bold and blue.
Species