Saprobic on dead plant parts. Sexual morph: Stromata irregularly scattered, conspicuous,
multi-loculate, superficial, with base slightly penetrating the epidermis, discoid to pulvinate,
reddish brown to black, scurfy surface, flat or slightly convex, composed of rather thin-walled,
reddish yellow cells of textura angularis, textura globosa or textura epidermoidea. Ascomata
perithecial, deeply imbedded in stromatic tissues, globose, with a long, periphysate, ostiolar neck.
Peridium thick, composed of several layers of reddish brown cells of textura angularis. Paraphyses
hypha-like, numerous, tapering towards the apex, not embedded in a gelatinous matrix. Asci 8-
spored, unitunicate, broad cylindrical, short pedicellate, apically rounded or truncate, with a J-,
discoid, refractive, apical ring. Ascospores bi-seriate, hyaline, 1-celled, ellipsoidal to cylindrical,
smooth-walled, lacking a mucilaginous sheath. Asexual morph: Undetermined.
Type family – Catabotryaceae Petr. ex M.E. Barr
Notes – Catabotryaceae was placed in Amplistromatales based on DNA sequence analyses
(Maharachchikumbura et al. 2015, 2016b). However, this family was later placed in
Sordariomycetes families incertae sedis based on phylogeny and molecular clock evidence
(Hongsanan et al. 2017). Catabotryaceae has only one strain representing its taxonomic placement
and more samples are needed to provide evidence for its ordinal status. Hyde et al. (2017a)
however suggested that a new order Catabotryales was needed to accommodate the monotypic
family Catabotryaceae based on divergent time estimations. Catabotryales is formally introduced
here for the single family Catabotryaceae. Catabotryales comprises saprobic species on dead leaves
and stems of tropical monocotyledons. This order is distinct from its sister orders in having
astromatic ascomata, broad cylindrical asci and ellipsoidal to cylindrical ascospores without a
mucilaginous sheath. The divergence time for Catabotryales has been estimated as 172 MYA.