Koralionastetales was introduced to accommodate the genera Pontogenia and Koralionastes (Campbell et al. 2009). Pontogenia was initially classified as Sordariomycetes, genera incertae sedis, while Koralionastes as Ascomycota, genera incertae sedis (Eriksson 2006). The order was established based on the monophyletic clade formed by three Koralionastes species and one Pontogeneia species in the LSU and SSU-based phylogeny (Campbell et al. 2009). Species of Koralionastetales are distinct from taxa belonging to other orders in Sordariomycetes by the formation of antheridia on their germinating ascospores (Zhang et al. 2006). Furthermore, all Koralionastetales species are restricted to marine habitats, in association with corals, sponges and algae (Campbell et al. 2009, Jones et al. 2019). The divergence time for Koralionastetales is estimated as 210.5 MYA (Fig. 2). Currently there is one family and two genera in this order (Hyde et al. 2020)