Members of Calosphaeriaceae and Pleurostomataceae are accommodated in Calosphaeriales
and are characterized by brown to black ascomata with reniform to clavate asci, phialidic
conidiophores and ellipsoid to allantoid conidia (Maharachchikumbura et al. 2016b, Huang et al.
2019). Some species are plant pathogens (Barr 1983, Damm et al. 2008, Trouillas et al. 2012). In
previous phylogenetic analyses, species of Calosphaeriales were closely related to Jobellisiales and
Togniniales (Maharachchikumbura et al. 2015, 2016b, Hongsanan et al. 2017). Calosphaeriales
comprises seven genera within the two families Calosphaeriaceae and Pleurostomataceae (Huang et
al. 2019). The monotypic genera, Kacosphaeria and Sulcatistroma are accepted in Calosphaeriales
based on the small, clavate asci with allantoid to ellipsoid, hyaline ascospores similar to
Calosphaeria (Huang et al. 2019). The divergence time for Calosphaeriales has been estimated as
160 MYA (Fig. 2). Currently there are two families and nine genera in this order (Hyde et al. 2020).