Lichenothelia D. Hawksw., Lichenologist 13(2): 142 (1981), MycoBank: MB 2855
Possible synonym: = Anzia Garov., Rev. Ist. Lomb., Milano, ser. 2 1: 558 (1868)
Thallus endolithic or epilithic, endokapylic or episubstratic, black, dispersed or continuous, areolate or not, rarely effigurate, or formed of superficial hyphae, branching or not, sometimes connecting stromata. Meristematic growth common. Sexual state: Ascomata perithecioid, ostiolate, with interascal filaments, the hymenial gel amyloid or not, or fertile stromata of various shapes, without an ostiole, the asci forming in pseudoparenchymatous locules, the interascal gel amyloid or not. Asci globose to broadly clavate, bitunicate, sometimes K/I+ bluish around the outer layers or apex of the ascus. Ascospores brown, 8 per ascus (but frequently few becoming mature), 1-septate to muriform (often variable), ellipsoid to subglobose, halonate or not, ornamented or not. Asexual state: Macroconidia black, originating from the surface of superficial hyphae, varying in shape and number of cells, common. Pycnidia, immersed in vegetative stromata, conidia hyaline, non-septate and rod-shaped (Henssen 1987) or unknown.

Type: Lichenothelia scopularia (Nyl.) D. Hawksw., Lichenologist 13(2): 147 (1981). MB 112068
Verrucaria scopularia Nyl., Not. Sällsk. Fauna et Fl. Fenn. Förh. 3: 85 (1861)

Fig. 61 Morphological and anatomical characters of Lichenothelia spp.: thallus of a L. scopularia . b L. calcarea. c L. intermixta. d L. tenuissima. e L. convexa; Transver sesection of ascoma, with asci and spores of f L. scopularia. g L. calcarea. h L. tenuissima. i L. intermixta. j k Asci of L. scopularia containing 8 mature spores. l, m Asci of L. tenuissima containing young (l) and mature (m) ascospores. n Ascus of L. intermixta containing 8 mature ascospores; ascospores of o L. calcarea. p–s L. scopularia. t–v L. intermixta. w L. tenuissima. Scale bars: a–c, e =300μm, d =1 mm, f–i =20μm, j–o =10μm, p–w=5μm. Photomicrographs were prepared in water. Specimens photographed are: Lichenothelia scopularia Obermayer 1252 (GZU000303500); L. calcarea Knudsen 11749 (62127 UCR1); L. intermixta Knudsen 13472 (UCR-220508); L. tenuissima Knudsen 10406 (UCR-197485); L. convexa Knudsen 12452 (1304KK64)

Fig. Morphological and anatomical characters of Lichenostigma spp. and Lichenostigma subgen. Lichenogramma spp.: a L. maureri. b L. rugosum. c L. epirupestre; transversal section of ascoma of d L. maureri and e, f L. epirupestre. g L. elongatum. h L. rouxii. i L. subradians; transversal section of ascoma of j, k L. elongatum and l L. subradians. Scale bars: a–c, g–i =0.4 mm, d–f, j–l =20μm. Photomicrographs were prepared in water. Specimens photographed are: Lichenostigma maureri on Pseudevernia furfuracea Hafellner 9886 (GZU000289898, paratype); L. rugosum on Diploschistes diacapsis Pérez-Ortega 1683; L. epirupestre on Pertusaria pertusa Pérez-Ortega 1433; L . elongatum on Lobothallia radiosa Hafellner 65534 (GZU000306021, paratype); L. rouxii on Squamarina cartilaginea Pérez-Ortega 1428; L. subradians on Acarospora sp. Hafellner 36841 (GZU000301218)