An order within the class Sordariomycetes, subclass Sordariomycetidae occurring as saprobes
or epiphytes on rotting or dead plants or fungi, and causing systemic mycotic infection in humans
with burns. Sexual morph: Ascomata perithecial, solitary to gregarious, carbonaceous, covered by
sulphureous hyphae, without periphyses. Peridium cephalothecoid. Ascogenous hyphae septate.
Asci 8-spored, unitunicate, evanescent, apedicellate, without an apical ring. Ascospores irregularly
arranged, brown, variously shaped, without germ pores. Asexual morph: Hyphomycetous.
Mycelium branched septate. Conidiophores long, stiffly upright, cylindrical, septate. Conidiogenous
cells phialidic, cylindrical. Conidia in chains, cylindrical, ovate or obovate, with or without an
apiculate or truncate base, 1-celled.
Type family – Cephalothecaceae Höhn.
Notes – Cephalothecaceae was introduced by Höhnel (1917c) and is characterized by
ascomata with a cephalothecoid peridium. The peridial cells form plate-like complexes made up of
radiating groups of cells, where each plate is separated by well-defined lines of dehiscence
(Malloch & Cain 1970). Albertiniella, Cephalotheca, Cryptendoxyla, Phialemonium and the newly
introduced Victoriomyces (Davolos et al. 2019) are presently placed in the family. Although the
placement of Cephalothecaceae is not stable within subclass Sordariomycetidae, it has high support
in the MCC tree (Hyde et al. 2017a, Hongsanan et al. 2017). Cephalothecaceae has a divergence
time at 154 MYA (Hyde et al. 2020), thus, here we raise Cephalothecaceae to ordinal status.